B.S., Northern Illinois University, 1964; Ph.D., University of Arizona, 1970; Postdoctoral Research Fellow, University of Michigan, 1970 - 1972.
Research Biochemist, University of California/San Francisco, 1972-1980; Visiting Professor, University of Milan, 1986, University of Konstanz, 1998; Member N.I.H. Physical Biochemistry Study Section, 1980-1984, 2002-2005; Member N.S.F. Predoctoral Fellowship Panel, 1985, 1987, 1988; Oversight Review Panel, Molecular Boiscience Division of N.S.F., 1985; Veterans Administration Basic Science Review Panel, 1987-90, Chairman, 1988-1990; Member, Cellular and Molecular Basis of Disease Reivew Committee N.I.G.M.S., 1991-1995.
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